Lichens of the Frankenhalvøya Peninsula, Northern of Barentsøya, Svalbard
DOI:
https://doi.org/10.24425/ppr.2021.138591Abstract
This paper contributes to the studies on the lichen diversity of Barentsøya. It covers 211 species, of which four (Buellia schaereri, Myriolecis zosterae var. palanderi, Rhizocarpon furfurosum, R. leptolepis) are reported for the first time for the Svalbard archipelago. Additionally, 84 of the species are reported for the first time for Barentsøya. Our study includes 2 subspecies as well, both new for Barentsøya. Thirty-six species (16.8% among the identified species) are rare in Svalbard, whereas more than two thirds (70.1% from identified in the Barentsøya) are relatively widespread species in Svalbard and the Arctic.
References
Ahti T., Jørgensen P.M., Kristinsson H., Moberg R., Søchting U. and Thor G. 2002. Nordic Lichen Flora Vol. 2. Physciaceae. TH-tryck AB, Uddevalla.
Ahti T., Jørgensen P.M., Kristinsson H., Moberg R., Søchting U. and Thor G. 2007. Nordic Lichen Flora Vol. 3. Cyanolichens. Mediaprint AB, Uddevalla.
Ahti T., Stenroos S. and Moberg R. 2013. Nordic Lichen Flora Vol. 5. Cladoniaceae. Zetterqvist tryckeri AB, Göteborg.
Alstrup V. and Olech M. 1993. Lichenicolous fungi from Spitsbergen. Polish Polar Research 14: 33–42.
Aptroot A. and Alstrup V. 1991. Lichens from Edgeøya, Svalbard. Graphis Scripta 3: 73–75.
Breuss O. 2017. Flechten aus Spitzbergen im Herbarium des Oberösterreichischen landesmuseums (LI), gesammelt 1975 von Werner Repetzky. Stapfia 107: 163–168.
Chesnokov S. and Konoreva L. 2015. Additions to the lichen biota of SE Siberia: records from the Stanovoye Nagor’e Highlands (Trans-Baikal Region, Russia). Polish Botanical Journal 60: 203–216.
Cooper E.J. and Wookey P.A. 2001. Field measurements of the growth rates of forage lichens, and the implications of grazing by Svalbard reindeer. Symbiosis 31: 173–186.
Czernyadjeva I.V., Ahti T., Boldina O.N., Chesnokov S.V., Davydov E.A., Doroshina G.Ya., Fedosov V.E., Khetagurov Kh.М., Konoreva L.A., Kotkova V.M., Kuzmina E.Yu., Lavrentiev M.V., Liksakova N.S., Nikolayev I.А., Popova N.N., Safronova T.V. and Shadrina S.N. 2020. New lichenological records. Novosti sistematiki nizshikh rastenii 54: 537–557.
Dobrysh A.A. 2003. Rhizocarpaceae М. Choisy ех Hafellner. In: Golubkova N.S. (eds.). Handbook of the lichens of Russia 8. Bacidiaceae, Catillariaceae, Lecanoraceae, Megalariaceae, Mycobilimbiaceae, Rhizocarpaceae, Trapeliaceae. Saint-Petersburg, Nauka: 198–238 (in Russian).
Elenkin A.A. 1907. Lichens collected on the Spitsbergen Islands by A. A. Byalynitskiy-Birulei and A. Bunge in 1899. Trudy Botanicheskago Muzeia Imperatorskoi Akademii Nauk 3: 60–63 (in Russian).
Elenkin A.A. and Savicz V.P. 1912. Lichens collected by I.V. Palibin in the voyage of the icebreaker Ermak in the Arctic Ocean in 1901. Acta Horti Petropolitani 32: 94–97 (in Russian).
Elvebakk A. 1984. Contributions to the lichen flora and ecology of Svalbard, Arctic Norway. The Bryologist 87: 308–313.
Elvebakk A. and Hertel H. 1996. A catalogue of Svalbard plants, fungi, algae and cyanobacteria. Part 6. Lichens. Norsk Polarinstitutt Skrifter 198: 271–359.
Elvebakk A. and Tønsberg T. 1992. Additions to the lichen flora of Svalbard. Graphis Scripta 3: 140–147.
Elvebakk A., Elven R. and Razzhivin V.Y. 1999. Delimitation, zonal and sectoral subdivision of the Arctic for the Panarctic Flora Project. In: Nordal I. and Razzhivin V.Y (eds.). The species concept in the High North – A panarctic flora initiative. The Norwegian Academy of Science and Letters, Oslo, Norway: 375–386.
Eurola S. 1968. Über die Fjeldheidevegetation in den Gebieten von Isfjorden und Hornsund in Westspitzbergen. Aquilo. Seria Botanica 7: 1–56.
Eurola S. 1971. Die Vegetation einer Sturzhalde (Sveagruva, Spitzbergen, 77°53′ n. Br.). Aquilo. Seria Botanica 10: 8–28.
Foucard T. 2001. Svenska skorplavar och svampar som växer på dem. Interpublish, Stockholm.
Foucard T., Moberg R. and Nordin A. 2002. Buellia. In: Ahti T., Jørgensen P.M., Kristinsson H., Moberg R., Søchting U. and Thor G. (eds.). Nordic Lichen Flora Vol. 2. Physciaceae. TH-tryck AB, Uddevalla: 11–25.
Fries T.M. 1860. Lichenes Arctoi. Ex Actis Regiae Societatis Scientiarum Upsaliensis Seriei III 3: 103–398.
Fries T.M. 1867. Lichenes Spitsbergenses. Kongl. Svenska Vetenskaps-Akademiens Handlingar 72: 1–53.
Gyelnik V. 1932. Enumeratio lichenum europaeorum novorum rariorumque. 1. Annales Mycologici 30: 442–459.
Hafellner J. 1982. Flechtenfunde am Bockfjord, Spitzbergen. Ergebnisse der Österreichischen Spitzbergen-Expedition 1979, I. Phyton 22: 23–50.
Hafellner J. 2010. Contributions to a revision of lichenized, phaeospored species of Polyblastia coll., mainly in Central European mountains. Bibliotheca Lichenologica 104: 117–141.
Halonen P., Myllys L., Velmala S. and Hyvärinen H. 2009. Gowardia (Parmeliaceae) – a new alectorioid lichen genus with two species. The Bryologist 112: 138–146.
Hansen E.S. 2009. Lichens from Johannes V. Jensen Land, N Greenland, the northernmost arctic land area. Willdenowia 39: 179–186.
Hertel H. 1977a. Bemerkenswerte Flechtenfunde aus dem Gebiet des Kongsfjordes und des Isfjordes (Spitzbergen). Herzogia 4: 367–401.
Hertel H. 1977b. Lecidea in der Arktis I. Mitteilungen der Botanischen Staatssammlung München 13: 337–352.
Hertel H. 1981. Lecidea in der Arktis II. Mitteilungen der Botanischen Staatssammlung München 17: 171–184.
Hertel H. 1991. Lecidea in der Arktis III (lecideoide Flechten, Lecanorales). Mitteilungen der Botanischen Staatssammlung München 30: 297–333.
Hertel H. and Ullrich H. 1976. Flechten von Amsterdamøya (Svalbard). Mitteilungen aus der Botanischen Staatssammlung München 12: 417–512.
Inoue T., Uchida M., Inoue M., Kaneko R., Kudoh S., Minami Y. and Kanda H. 2019. Vegetation data of high Arctic lichens on Austre Brøggerbreen glacier foreland, Ny-Ålesund, Svalbard, in 1994. Polar Data Journal 3: 1–11.
Jørgensen P.M. and Zhurbenko M. 2002. Two new, remarkable, Arctic species in the lichen genus Fuscopannaria (Pannariaceae, lichenized Ascomycetes). The Bryologist 105: 465–469.
Kondratyuk S.Y., Lőkös L., Farkas E., Kärnefelt I., Thell A., Yamamoto Y. and Hur J.-S. 2020. Three new genera of the Teloschistaceae proved by three gene phylogeny. Acta Botanica Hungarica 62: 109–136.
Konoreva L.A. 2011. A contribution to the lichens of vicinity of the former Pyramiden settlement (Spitsbergen Island, Svalbard Archipelago). Novosti sistematiki nizshikh rastenii 45: 183–193 (in Russian).
Konoreva L.A. and Chesnokov S.V. 2018. Lichens of vicinity of the former township Kolsbey (West Spitsbergen). Bulletin of the Kola Scientific Center of the Russian Academy of Sciences 3: 31–42 (in Russian).
Konoreva L.A. and Chesnokov S.V. 2021. Lichens of Innvika Bay, Prins Oscars Land (Nordaustlandet, Svalbard). Czech Polar Reports 11: 253–269.
Konoreva L.A., Kholod S.S., Chesnokov S.V. and Zhurbenko M.P. 2019a. Lichens of Franz Josef Land archipelago. Polish Polar Research 40: 139–170.
Konoreva L., Kozhin M., Chesnokov S. and Hong S.G. 2019b. Lichens and vascular plants in Duvefjorden area on Nordaustlandet, Svalbard. Czech Polar Reports 9: 182–199.
Kristinsson H., Zhurbenko M.P. and Hansen E.S. 2010. Panarctic checklist of lichens and lichenicolous fungi. CAFF Technical Report 20: 1–120.
Krzewicka B. and Maciejowski W. 2008. Lichen species from the northeastern shore of Sørkapp Land (Svalbard). Polar Biology 31: 1319–1324.
Lumbsch H.T. and Huhndorf S.M. 2010. Myconet Volume 14. Part One. Outline of Ascomycota—2009. Part Two. Notes on Ascomycete Systematics. Nos. 4751–5113. Fieldiana Life and Earth Sciences 1: 1–64.
Lynge B. 1926. Lichens from Bear Island (Bjørnøya) collected by Norwegian and Swedish expeditions, chiefly by Th. M. Fries during the Swedish polar expedition of 1868. Resultater av de Norske Statsunderstøttede Spitsbergen Ekspeditioner 1: 1–78.
Lynge B. 1938. Lichens from the west and north coasts of Spitsbergen and the North-East Land collected by numerous expeditions. 1. The macrolichens. Skrifter Norsk videnskaps-akademi i Oslo. 1. Mathematisk-naturvidenskabelig klasse 6: 1–136.
Lynge B. 1940. Et bidrag til Spitsbergens lavflora. Laver samlet av Emil Hadač, fortrinsvis i Sassenområdet, sommeren 1939. Skrifter om Svalbard og Ishavet 79: 1–22.
Maciejowski W., Osyczka P., Smykla J., Ziaja W., Ostafin K. and Krzewicka B. 2018. Diversity and distribution of lichens in recently deglaciated areas of southeastern Spitsbergen. Acta Societatis Botanicorum Poloniae 87: 3596.
Magnusson A.H. 1935. The lichen-genus Acarospora in Greenland and Spitsbergen. Nyt Magazin for Naturvidenskaberne 75: 221–241.
Magnusson A.H. 1944. Studies in the ferruginea-group of the genus Caloplaca. Kungliga Vetenskaps- och Vitterhets-samhälles Handlingar, Sjätte Följden, serie B 3: 1–71.
Magnusson A.H. 1950. On some species of Blastenia and Caloplaca with black apothecia. Botaniska Notiser 1950: 369–386.
Makryi T.V. 2002. To the lichen flora of the Stanovoye Nagorye Uplands (Baikalian Siberia). I. The epilithic lichens of the Kodar Range. Turczaninowia 5: 47–67 (in Russian).
Mattick F. 1949. Die Flechten Spitzbergens. Polarforschung 19: 261–273.
Mayrhofer H. 1984. Die saxicolen Arten der Flechtengattung Rinodina und Rinodinella in der alten Welt. Journal of the Hattori Botanical Laboratory 55: 327–493.
Moberg R., Tibell S. and Tibell L. 2017. Nordic Lichen Flora Vol. 6. Verrucariaceae 1. Billes Tryckeri AB, Mölndal.
Nimis P.L. 1985. Structure and floristic composition of a high arctic tundra: Ny Ålesund (Svalbard archipelago). Inter-Nord 17: 47–58.
Nowak J. 1965. The lichens from Hornsund (S.W.-Spitsbergen) collected during the Polish polar expeditions in 1957 and 1958. Fragmenta Floristica et Geobotanica 11: 171–190.
Olech M. 1987. Materials to the lichen flora of Hornsund (SW Spitsbergen). Zeszyty Naukowe Uniwersytetu Jagiellońskiego, Prace Botaniczne 15: 165–168.
Olech M. 1990. Lichens of the NW Sørkapp Land (Spitsbergen). Zeszyty Naukowe Uniwersytetu Jagiellońskiego, Prace Botaniczne 21: 197–210.
Olech M. and Alstrup V. 1989. Lichens new to Spitsbergen. Graphis Scripta 2: 146–148.
Osyczka P. 2006. The lichen genus Cladonia (Cladoniaceae, lichenized Ascomycota) from Spitsbergen. Polish Polar Research 27: 207–242.
Osyczka P. and Węgrzyn M. 2008. Lichens on lignum in the coastal regions of western Spitsbergen (Svalbard). Biologia 63: 1069–1072.
Øvstedal D.O., Tønsberg T. and Elvebakk A. 2009. The lichen flora of Svalbard. Sommerfeltia 33: 1–393.
Paulson R. 1928. Lichens of Spitsbergen and North-East Land. Journal of Botany 66: 249–253.
Redchenko O., Košnar J. and Gloser J. 2010. A contribution to lichen biota of the central part of Spitsbergen, Svalbard Archipelago. Polish Polar Research 31: 159–168.
Saag L., Saag A. and Randlane T. 2009. World survey of the genus Lepraria (Stereocaulaceae, lichenized Ascomycota). The Lichenologist 41: 25–60.
Schuhwerk F. 1992. Die Berücksichtigung der Ökologie in der Lichenometrie: Datierung mit Sukzessionsstadien von Flechtengesellschaften. In: Blümel W.D. (ed.). Geowissenschaftliche Spitzbergen-Expedition 1990 und 1991 "Stofftransporte Land-Meer in polaren Geosystemen". Stuttgarter Geographische Studien 117: 161–175.
Sedelnikova N.V. 2013. Species diversity of lichen biota of the Altai-Sayan ecological region. Plant Life of Asian Russia 2: 12–54 (in Russian).
Singh S.M. and Ravindra R. 2013. Chapter 7. Impact of Climate Change on Lichen and Moss Communities in Ny-Ålesund, Arctic: Some Preliminary Observations. In: Sinha R. and Ravindra R. (eds.). Earth System Processes and Disaster Management. Society of Earth Scientists Series 1, Springer-Verlag Berlin Heidelberg: 93–100.
Svalbard Lichens Database. 2008. https://nhm2.uio.no/botanisk/nxd/sval_L/sld_e.htm
Śliwa L. 2007. A revision of the Lecanora dispersa complex in North America. Polish Botanical Journal 52: 1–70.
Smith C.W., Aptroot A., Coppins B.J., Fletcher A., Gilbert O.L., James P.W. and Wolseley P.A. 2009. The lichens of Great Britain and Ireland. The British Lichen Society, London.
Søchting U. 1989. Lignicolous species of the lichen genus Caloplaca from Svalbard. Opera Botanica 100: 241–257.
Søchting U. 1992. On the identity and distribution of some Nordic Caloplaca species. Graphis Scripta 4: 91–92.
Søchting U. and Olech M. 1995. The lichen genus Caloplaca in polar regions. The Lichenologist 27: 463–471.
Søchting U., Balschmidt Lorentsen L. and Arup U. 2008. The lichen genus Caloplaca (Ascomycota, Lecanoromycetes) on Svalbard. Notes and additions. Nova Hedwigia 87: 69–96.
Šoun J., Vondrák J., Søchting U., Hrouzek P., Khodosovtsev A. and Arup U. 2011. Taxonomy and phylogeny of the Caloplaca cerina group in Europe. The Lichenologist 43: 113–135.
Summerhayes V.S. and Elton C.S. 1928. Further contributions to the ecology of Spitsbergen. Journal of Ecology 16: 193–268.
Thell A. and Moberg R. 2011. Nordic Lichen Flora Vol. 4. Parmeliaceae. Zetterqvist tryckeri, Göteborg.
Urbanavichene I.N. and Koroleva N.E. 2008. Lichens. In: Koroleva N.E., Konstantinova N.A., Belkina O.A., Davydov D.A., Likhachev A.Yu., Savchenko A.N. and Urbanavichene I.N. (eds.). Flora of Vegetation of Grønfjord area (Spitsbergen archipelago). K&M, Apatity: 81–92 (in Russian).
Urbanavichus G.P. 2010. A checklist of the lichen flora of Russia. St. Petersburg, Nauka (in Russian).
Urbanavichus G.P. and Urbanavichene I.N. 2004. Lichens. In: Afonina O.M. and Golubkova N.S. (eds.). The present-day state of biological diversity within protected areas in Russia. Issue 3. Lichens and Bryophytes. IUCN – The World Conservation Union, Moscow: 5–235 (in Russian).
Vainio E.A. 1905. Lichenes expeditionis G. Amdrup (1898-1902). Meddelelser om Grønland 30: 125–141.
Vězda J. 1967. Flechtensystematische Studien VI. Die Gattung Sagiolechia Massal. Folia Geobotanica et Phytotaxonomica 2: 383–396.
Vitikainen O. 1994. Taxonomic revision of Peltigera (lichenized Ascomycotina) in Europe. Annales Botanici Fennici 152: 1–96.
Vitikainen O. 2007. Peltigeraceae. In: Ahti T., Jørgensen P.M., Kristinsson H., Moberg R., Søchting U. and Thor G. (eds.). Nordic Lichen Flora 3. Cyanolichens. The Nordic Lichen Society, Museum of Evolution, Uppsala University, Uddevalla: 113–131.
Węgrzyn M., Wietrzyk P., Adamska E. and Nicia P. 2015. New records of driftwood lichens in the Kaffiøyra Plain (NW Spitsbergen, Svalbard). Polish Polar Research 36: 189–195.
Węgrzyn M.H., Wietrzyk-Pełka P., Galanty A., Cykowska-Marzencka B. and Sundset M.A. 2019. Incomplete degradation of lichen usnic acid and atranorin in Svalbard reindeer (Rangifer tarandus platyrhynchus). Polar Research 38: 3375.
Westberg M. 2007. Candelariella (Candelariaceae) in western United States and northern Mexico: the polysporous species. The Bryologist 110: 375–390.
Westberg M., Moberg R., Myrdal M., Nordin A. and Ekman S. 2021. Santesson’s Checklist of Fennoscandian Lichen-Forming and Lichenicolous Fungi. Uppsala University, Museum of Evolution.
Wietrzyk P., Węgrzyn M. and Lisowska M. 2016. Vegetation diversity and selected abiotic factors influencing the primary succession process on the foreland of Gåsbreen, Svalbard. Polish Polar Research 37: 493–509.
Wietrzyk P., Węgrzyn M. and Lisowska M. 2017. Lichen diversity on glacier moraines in Svalbard. Cryptogamie, Mycologie 38: 67–80.
Wietrzyk-Pełka P., Otte V., Węgrzyn M.H. and Olech M. 2018. From barren substrate to mature tundra – lichen colonization in the forelands of Svalbard glaciers. Acta Societatis Botanicorum Poloniae 87: 3599.
Wietrzyk-Pełka P., Rola K., Patchett A., Szymański W., Węgrzyn M.H. and Björk R.G. 2021. Patterns and drivers of cryptogam and vascular plant diversity in glacier forelands. Science of The Total Environment 770: 144793.
Zhurbenko M.P. 2010. Lichenicolous fungi and lichens growing on Stereocaulon from the Holarctic, with a key to the known species. Opuscula Philolichenum 8: 9–39.
Zhurbenko M.P. and Brackel W.V. 2013. Checklist of lichenicolous fungi and lichenicolous lichens of Svalbard, including new species, new records and revisions. Herzogia 26: 323–359.
Ziaja W., Maciejowski W. and Ostafin K. 2009. Coastal Landscape Dynamics in NE Sørkapp Land (SE Spitsbergen), 1900–2005. Ambio 38: 201–208.
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